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'A # B' means 'A is the brother of B'. 'A @ B' means 'A is the daughter of B'. 'A & B' means 'A is the husband of B'. 'A % B' means 'A is the wife of B'. If N @ O # P % Q # R & S, then how is O related to R?

A Father's brother

B Brother's wife

C Sister-in-law's brother

D Mother's brother

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