Sentences/segments of a paragraph are given below in jumbled order. Arrange them in the correct order to form a meaningful and coherent paragraph.  
A. this raises concerns about the long-term sustainability of logging in the tropics 
B. trees of recovering tropical forests were found to have tougher leaves 
C. and are pushing the nutrient content towards ecological limits  
D. both essential for plant and tree growth, than trees of old-growth forests   E. this suggests that multiple cycles of logging and recovery irreversibly remove phosphorus from the forest system  F. with lower concentrations of nutrients such as phosphorus and nitrogen