Question

In a certain code language, 'A # B' means 'A is the sister of B', 'A @ B' means 'A is the son of B', 'A & B' means 'A is the wife of B' and 'A % B' means 'A is the father of B'. How is Q related to R, if 'U # V % Q @ P # S & T @ R'?

A Son
B Nephew
C Grandfather
D Grandson
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