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Net Primary Productivity (NPP) is the energy or biomass that remains after autotrophs (producers) have used some of it for respiration (R). It represents the energy available for the next trophic level, i.e., herbivores (primary consumers). It is calculated as: NPP=GPP−R where GPP (Gross Primary Productivity) is the total energy fixed by autotrophs through photosynthesis, and R is the energy used in respiration.
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